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Because of the strict scientific definition we do have to exclude humans for all of Doyle's reasons. Apex predators are the king of their habitat/biome/environment.Because man can adapt to his surroundings with his tools (merchanical as well as menta) in any of the earths' environment, and for that fact even the moons environment he is king. Lions in their area, white sharks in theirs and the list goes on are at the top of the heap in their own back yard. Even put a Betta fish in an aquarium full of guppies and the king will emerge.