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They always start with a standard caliber or standard case.  Such as the 30-06 makes into the 30-06 Ackley improved.  The 270 into the 270 Ackley improved and the 243 Remington into the 243 Ackly improved. 


Basically a tapered case is blown out straight and the shoulder angles made sharper to give the case both more grip on the chamber walls (straighter cases do that) and a little more powder capacity.  They are also loaded somewhat hotter (more pressure than most standard cartridges).


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